Wednesday 4 July 2012

Chapter I: The Inner Workings of HIV


As far as the phrasing goes: Know your enemy and know your enemy well in order to win the day. It's a  little cliche but a touche ^^
In order to fully fathom the premise of HIV treatment or possibly cure, we must first be able to understand how this little midget works



HIV virion

Entry (This virus do know how to make an entrance^^): [1]


HIV infection is specific to certain human immune cells (helper T-cell) due to the presence of the protein CD4 (exclusive to helper T cells) which acts as a receptor that allow the virus to enter the cell; other than the CD4 receptor, a secondary receptor CCR5* (Chemokine Co-receptor) is also needed. After the virus attaches itself to these cell-surface receptors, it then fuses the viral membrane and the cell membrane and inject the viral particles into the cell while leaving its membrane protein behind.


Reverse Transcription and Integration


After the virus make its way into the cell, its matrix and capsid protein break down to release the viral genetic material. Then the enzyme, reverse transcriptase* that the virus brings along with it starts acting on the viral mRNA to transcribe a complementary DNA or cDNA. Now, another enzyme that the virus brought along, integrase* would bring the viral DNA into the cellular nucleus and integrate that piece of viral DNA into the human DNA, which dawns upon the start of a life-long infection as the virus take advantage of the cellular machinery to create more copies of itself and infect neighboring cells. Once integrated, the HIV DNA is known as provirus.

Transcription and Translation


HIV provirus may lie dormant within a cell for a long time. But when the cell becomes activated, it treats HIV genes in much the same way as human genes. First it converts them into messenger RNA (using human enzymes). Then the messenger RNA is transported outside the nucleus, and is used as a blueprint for producing new HIV proteins and enzymes.

[2]Interesting tidbits: While HIV only has 9 genes, it’s able to take advantage of mRNA slippage and a change in reading frame to generate different protein from the same message (Programmed frameshifting in biosynthesis of HIV protein). A single mRNA, designated gag-pol, encodes for two polyproteins that overlap by about 200 nucleotides and are in different reading frames. The gag polyprotein is translated from the initiation codon to an in-frame termination codon near the gag-pol junction.; gag polyprotein is then cleaved to generate several structural proteins of the virus. However about 5% of the time a one-nucleotide frameshift occurs within the overlapping segment of the mRNA and the termination codon is bypassed because it is no longer in the reading frame. A gag-pol fusion polyprotein is produced; proteolytic cleavage of the pol polyprotein produces the viral reverse transcriptase and other proteins needed in virus replication.



Assembly, Budding and Maturation

Among the strands of messenger RNA produced by the cell are complete copies of HIV genetic material. These gather together with newly made HIV proteins and enzymes to form new viral particles. The HIV particles are then released or 'bud' from the cell. The enzyme protease* plays a vital role at this stage of the HIV life cycle by chopping up long strands of protein into smaller pieces, which are used to construct mature viral cores. 

The newly matured HIV particles are ready to infect another cell and begin the replication process all over again. In this way the virus quickly spreads through the human body. And once a person is infected, they can pass HIV on to others in their bodily fluids.


HIV budding from an infected cell

If you are too lazy to read the whole process, here's an olive branch ^^:


References:
[2] Glitz, D.(2006). Textbook of Biochemistry With Clinical Correlations Sixth Edition. Canada: Wiley-Liss.

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